| Haldan K. Hartline  AKA Haldan Keffer Hartline
Born: 22-Dec-1903
Birthplace: Bloomsburg, PA
Died: 17-Mar-1983
Location of death: Fallston, MD
Cause of death: Natural Causes
Remains: Buried, Bloomsburg, PA
Gender: Male
Race or Ethnicity: White
Occupation: Scientist, Doctor Nationality: United States
Executive summary: Physiology of the retina American physiologist Haldan K. Hartline spent decades studying the optic nerves of frogs and horseshoe crabs. Working with Columbia University physiologist Clarence H. Graham, he was the first scientist to isolate and record of the activity of a single optic nerve fiber. He later showed that different fibers in the optic nerves respond to light in different ways. For his research on cellular-level electrical activity within the eye, Hartline was awarded the Nobel Prize in Medicine in 1967, shared with Ragnar Granit and George Wald. He always preferred to be called by his middle name, Keffer, and upon winning his Nobel honors, Dr Hartline said he was "not very comfortable about all the fuss." Father: Daniel Schollenberger Hartline (biologist, b. 19-Sep-1866, d. 1946)
Mother: Harriet Franklin Keffer Hartline (teacher, b. 1864, m. 26-Aug-1897, d. 1946)
Wife: Elizabeth Kraus Hartline ("Beth", psychologist, b. 17-Sep-1909, m. 11-Apr-1936)
Son: Daniel Keffer Hartline (neurobiologist, b. 1-Dec-1939)
Son: Peter Haldan Hartline (biophysicist, b. Jan-1942)
Son: Frederick Flanders Hartline (physicist, b. 13-Feb-1945)
Scholar: Biology, Woods Hole Oceanographic Institution (1921-22)
University: BS Biology, Lafayette College (1923)
Medical School: MD, Johns Hopkins University (1927)
Scholar: Physics, Johns Hopkins University (1927-29)
Scholar: Physics, University of Pennsylvania (1929-30)
Scholar: University of Leipzig (1930-31)
Scholar: University of Munich (1931)
Teacher: Physics, University of Pennsylvania (1931-40, 1941-49)
Professor: Physiology, Cornell University (1940-41)
Professor: Biophysics, Johns Hopkins University (1949-53)
Professor: Neurophysiology, Rockefeller University (1953-83)
Nobel Prize for Medicine with Ragnar Granit and George Wald (1967)
American Academy of Arts and Sciences
American Philosophical Society
American Physiological Society
National Academy of Sciences
National Research Council Fellowship (1927)
Optical Society of America
Royal Society
Author of books:
Studies on Excitation and Inhibition in the Retina (1974)
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